Nature lovers held a vigil for a famous female grizzly bear that delighted visitors for decades in Wyoming’s Grand Teton National Park after she was struck and killed by a vehicle

She was a Yellowstone icon with more than 20 cubs and grand cubs, and the grizzly bear population will surely suffer from her loss."

A female grizzly bear who was adored by thousands of visitors to Grand Teton National Park was struck and killed on Tuesday, park officials announced.

During her nearly three decades roaming the snowy wilderness of the Teton Range, Grizzly No. 399 became a beloved mama bear with millions of parkgoers following her yearly adventures and her ever-growing family tree.

During her nearly three decades roaming the snowy wilderness of the Teton Range, Grizzly No. 399 became a beloved mama bear with millions of parkgoers following her yearly adventures and

Named for the tag affixed by researchers to her ear, No. 399 was the oldest-known reproducing female grizzly in the greater Yellowstone ecosystem